John H. Willis

Professor of Biology

3314 French Science Center, Science Drive, Durham, NC 27708
Campus Box: 
Box 90338, Department of Biology, Durham, NC 27708-0338
(919) 660-7340
We conduct research on broad issues in evolutionary genetics, and we are currently addressing questions relating to the evolution of adaptation, reproductive isolation, breeding systems, inbreeding depression, and floral traits in natural plant populations. Please see our lab web page for more information.


  • Ph.D., The University of Chicago 1991

Modliszewski, Jennifer L., and John H. Willis. “Allotetraploid Mimulus sookensis are highly interfertile despite independent origins.Molecular Ecology 21, no. 21 (November 2012): 5280–98. Full Text

Mojica, Julius P., Young Wha Lee, John H. Willis, and John K. Kelly. “Spatially and temporally varying selection on intrapopulation quantitative trait loci for a life history trade-off in Mimulus guttatus.Molecular Ecology 21, no. 15 (August 2012): 3718–28. Full Text

Strasburg, Jared L., Natasha A. Sherman, Kevin M. Wright, Leonie C. Moyle, John H. Willis, and Loren H. Rieseberg. “What can patterns of differentiation across plant genomes tell us about adaptation and speciation?Philosophical Transactions of the Royal Society of London. Series B, Biological Sciences 367, no. 1587 (February 2012): 364–73. Full Text

Lowry, David B., Calvin C. Sheng, Zhirui Zhu, Thomas E. Juenger, Brett Lahner, David E. Salt, and John H. Willis. “Mapping of ionomic traits in Mimulus guttatus reveals Mo and Cd QTLs that colocalize with MOT1 homologues.Plos One 7, no. 1 (January 24, 2012): e30730. Full Text

Lowry, David B., Calvin C. Sheng, Jesse R. Lasky, and John H. Willis. “Five anthocyanin polymorphisms are associated with an R2R3-MYB cluster in Mimulus guttatus (Phrymaceae).American Journal of Botany 99, no. 1 (January 2012): 82–91. Full Text

Moran, Emily V., John Willis, and James S. Clark. “Genetic evidence for hybridization in red oaks (Quercus sect. Lobatae, Fagaceae).American Journal of Botany 99, no. 1 (January 2012): 92–100. Full Text

Benedict, B. G., J. L. Modliszewski, A. L. Sweigart, N. H. Martin, F. R. Ganders, and J. H. Willis. “Mimulus sookensis (Phyrmaceae), a new allotetraploid species derived from Mimulus guttatus and Mimulus nasutus.Madrono 59 (2012): 29–43.

Scoville, Alison G., Young Wha Lee, John H. Willis, and John K. Kelly. “Explaining the heritability of an ecologically significant trait in terms of individual quantitative trait loci.Biology Letters 7, no. 6 (December 2011): 896–98. Full Text

Magwene, Paul M., John H. Willis, and John K. Kelly. “The statistics of bulk segregant analysis using next generation sequencing.Plos Comput Biol 7, no. 11 (November 2011): e1002255. Full Text