John H. Willis

Professor of Biology

3314 French Science Center, Science Drive, Durham, NC 27708
Campus Box: 
Box 90338, Department of Biology, Durham, NC 27708-0338
(919) 660-7340
We conduct research on broad issues in evolutionary genetics, and we are currently addressing questions relating to the evolution of adaptation, reproductive isolation, breeding systems, inbreeding depression, and floral traits in natural plant populations. Please see our lab web page for more information.


  • Ph.D., The University of Chicago 1991

Fishman, L., A. J. Kelly, and J. H. Willis. “Minor QTLs underlie floral characters associated with mating system divergence in Mimulus.” Evolution 56 (2002): 2138–55.

Fishman, L., A. J. Kelly, E. Morgan, and J. H. Willis. “A genetic map in the Mimulus guttatus species complex reveals transmission ratio distortion due to heterospecific interactions.Genetics 159, no. 4 (December 2001): 1701–16.

Fishman, L., and J. H. Willis. “Evidence for Dobzhansky-Muller incompatibilites contributing to the sterility of hybrids between Mimulus guttatus and M. nasutus.Evolution; International Journal of Organic Evolution 55, no. 10 (October 2001): 1932–42. Full Text

Kelly, J. K., and J. H. Willis. “Deleterious mutations and genetic variation for flower size in Mimulus guttatus.Evolution; International Journal of Organic Evolution 55, no. 5 (May 2001): 937–42.[0937:dmagvf];2. Full Text

Willis, J. H. “The role of genes of large effect on inbreeding depression in Mimulus guttatus.” Evolution 53, no. 6 (December 1, 1999): 1678–91.

Schultz, S. T., M. Lynch, and J. H. Willis. “Spontaneous deleterious mutation in Arabidopsis thaliana.Proceedings of the National Academy of Sciences of the United States of America 96, no. 20 (September 1999): 11393–98. Full Text

Lynch, M., J. Blanchard, D. Houle, T. Kibota, S. Schultz, L. Vassilieva, and J. Willis. “Perspective: Spontaneous deleterious mutation.” Evolution 53, no. 3 (January 1, 1999): 645–63. Full Text

Sweigart, A., K. Karoly, A. Jones, and J. H. Willis. “The distribution of individual inbreeding coefficients and pairwise relatedness in a population of Mimulus guttatus.” Heredity 83, no. 5 (January 1, 1999): 625–32. Full Text